Not quite the same situation as described in the article, but it is still possible for the kernel to swap memory in and out of disk even without a swap file/partition. Memory used for storing executable binaries is allowed to be moved out of memory, as a copy of it lives on disk. This means you can still encounter memory thrashing (and thus system unresponsiveness) under low memory situations.
> Memory used for storing executable binaries is allowed to be moved out of memory, as a copy of it lives on disk.
On Linux, this is not necessarily the case, as you can change the file on disk while the executable is running. I don't know if Linux just keeps executable code in memory all the time, or if it is smart enough to detect whether a copy of executable pages still lives on disk.